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Granting that it is an older post, two aspects stand out to me. The first is that females were swapped in and out without giving them further time to establish a series of reproductive efforts per female. Successive efforts without withdrawing females alone may have solved or improved things. The other is that there is a formulaic aspect as well as a personal skill aspect to incubation aside from the variability of the status/condition of the eggs one is incubating. While I do not work with Colubrid breeding, I have bred and do breed a lot of things. Giving females time to "season" or find a rhythm can be important. Learning the nuances of individual females can be important. Getting a base formula (or method or recipe) that is generally successful is important. Then getting an intuitive feel for things can further squeeze out positive increments of productivity that a rote formula alone does not quite achieve. This last part has made an especially meaningful and rewarding impact on my experiences and results as a reptile breeder.
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