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"...Could they have been a harmless mite in the substrate which appeared after her death?"
No, "harmless mites" wouldn't be feeding on her as these are. I'm sorry for your loss, more so for your snake, but you missed the mites on this snake when you got her, & using a substrate they didn't show up on helped them hide until they did what they always do without treatment: their population exploded exponentially in a short period of time & killed your snake by sucking her blood. It's a tough way to learn this lesson. I'd be putting all my other snakes on white paper towels for a while, immediately giving them supervised soaks in mildly soapy water (Ivory) & quickly investing in some "PAM" (Provent-A-Mite), as mites travel to other cages and it would be a miracle if your other snakes don't have them too. You can accidentally transport them from one cage to another on your clothes, & you should assume that you did.
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